# Tracking and modelling native woodland expansion in the Scottish Highlands using LiDAR

*2026-09-01 — paper*

Authors: Sudina Thapa, Aland H. Y. Chan, Philippa R. Gullett, William R. M. Flynn, Liam Wakefield, Mark H. Hancock, Anil Madhavapeddy, David A. Coomes


Sustained culling of deer in parts of the Scottish Highlands is resulting in
natural recolonisation of open landscapes by woodlands. Tracking these changes
and furthering our understanding of the drivers of expansion is challenging due
to the limited scale of field‐collected data.
                
We developed a new pipeline to automatically segment and map small regenerating
trees with airborne LiDAR data and applied it to a 569 km2 study site over
Cairngorms Connect -- the largest contiguous restoration partnership in the
United Kingdom.

The results were validated by young trees accurately geolocated and measured in
the field. We then used the map of regenerating trees to build a Hurdle Model
and assessed the influence of topographical and biophysical factors on
regeneration success.
                  
Results show that clusters of regenerating trees are detectable once they
outgrow neighbouring shrubs (up to 1.1 m in height). Detection rates reached
83% and \>95% for tree clusters 1.4–1.6 m and \>1.8 m in height, respectively.
The algorithm also gave good estimates of the heights (R2=0.94) and canopy
cover (R2=0.90) of young trees, whilst keeping commission errors low (0.019%).
                  
Deer control has led to large‐scale natural forest colonisation in Cairngorms
Connect. The new pipeline delineated 2.65 million young trees (\<5 m in height)
in the study area. It is estimated that 5325 ha of previously non‐forested land
now records sapling density \>100ha. Geospatially, proximity to seed sources,
elevation, habitat type and deer density all strongly affected tree
establishment and sapling density.
                  
Forest restoration projects should utilise the new LiDAR‐based pipelines to
effectively map young trees, assess whether targets are met, and plan further
action. Notably, the Scottish Government is now commissioning a national
airborne LiDAR scan, providing a unique opportunity to upscale our algorithms
to create national woodland expansion databases.


DOI: 10.1111/1365-2664.70561
Classification: full
Venue: Journal of Applied Ecology
Volume: 63
Issue: 9
URL: https://besjournals.onlinelibrary.wiley.com/doi/10.1111/1365-2664.70561

## Related

- [Mapping millions of little trees across the Cairngorms with airborne LiDAR](https://anil.recoil.org/notes/cairngorms-woodland-expansion) (note, 2026-09-24)

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Canonical: https://anil.recoil.org/papers/2026-cairngorms-expansion
Type: paper
License: CC BY 4.0 <https://creativecommons.org/licenses/by/4.0/>
Tags: journal
